European equities traded in the US as American depositary receipts were declining late Wednesday morning, falling 0.58% to 1,880.17 on the S&P Europe Select ADR Index.

From continental Europe, the gainers were led by telecommunications company Nokia (NOK) and petroleum refiner Equinor (EQNR), which rose 2.1% and 1.3%, respectively. They were followed by oil and gas company Eni (E) and biopharmaceutical company argenx (ARGX), which were up 1% and 0.5%, respectively.

The decliners from continental Europe were led by internet ad firm Criteo (CRTO) and biopharmaceutical company Cellectis (CLLS), which fell 7% and 4.7%, respectively. They were followed by biotech firm Evaxion (EVAX) and software company SAP (SAP), which were down 4.6% and 4.5%, respectively.

The gainers from the UK were led by Mereo BioPharma Group (MREO), which advanced 5.6%. It was followed by pharmaceutical company Silence Therapeutics (SLN) and medical device maker Smith & Nephew (SNN), which increased 4.8% and 2%, respectively.

The decliners from the UK and Ireland were led by communications company WPP (WPP) and software company Endava (DAVA), which dropped 4.5% and 4.4%, respectively. They were followed by lender Barclays (BCS) and biotech firm Autolus Therapeutics (AUTL), which lost 2.4% and 1.7%, respectively.